Puss in Boots discovers that his passion for adventure and his contempt for danger ended up costing him dearly: he used up eight of his nine lives, and lost count in the process. In order to get back on his feet, our hairy hero literally embarks on the quest of his life.
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He embarks on an epic adventure to the confines of the Dark Forest in order to unearth the mythical Wishing Star, the only one capable of restoring his lost lives. But when you only have one left, you have to know how to keep a low profile, be careful and ask for help. This is how he turns to his former partner and best enemy of all time: the bewitching Kitty Velvet Paws. Puss in Boots and the beautiful Kitty will be helped in their quest, against their will, by Perro, a wandering and scaly lout with a strong tongue and an unalterable good humor.