In a world where beings of fire, water, air, and earth exist, Ember Lumen (Leah Lewis) is a first generation fire elemental and daughter of immigrants Bernie and Cinder (Ronnie del Carmen and Shila Ommi respectively) who emigrated to Element City facing prejudice and systemic segregation to start a bodega, called The Fireplace, that is the unofficial hub of Firetown which Bernie hopes Ember will someday inherit. Ember routinely struggles to keep her temper with customers and during one such episode inadvertently damages an old water pipe which pulls in city inspector Wade Ripple (Mamoudou Athie) from the other side of town and must reluctantly issue several citations on the bodega.
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After learning of Ember's struggles with trying to takeover The Fireplace from her ailing father, Wade feeling guilty over it tries to help Ember save The Fireplace as well as find out why there is water still running through Firetown when it shouldn't and eventually a romance blooms in spite of the historic incompatibility with fire and water.